University of North Carolina Wilmington
Myrtle Grove 1329, Deloach 117
Tel: 910 962-7406
Hampton Beach, NH
My research focuses on the evolution and paleoecology of Coastal Plain molluscs. I am especially interested in the evolutionary process and the factors that control it, and I believe that the fossil record contains data essential for deciphering this process. My initial studies, a test of the hypothesis of punctuated equilibrium, investigated tempo and mode of evolution of the Miocene mollusc fauna of Maryland. More recent and current work focuses on evolutionary paleoecology, with a particular interest in the role of ecological factors, such as predation, in evolution. A major thrust of this work has involved testing Vermeij’s hypothesis of escalation, which proposes that biological hazards such as predation, along with adaptation to those hazards, have increased through geological time. Thor Hansen (Western Washington University) and I have compiled a database on predation by shell-drilling naticid gastropods in the Gulf and Atlantic Coastal Plain; our collections now include >150,000 mollusc specimens of Cretaceous through Pleistocene age. We have used these data to test a variety of hypotheses about the dynamics and evolutionary history of predator-prey interactions, predator-prey coevolution and escalation, the ecology of mass extinctions and recoveries of mollusc faunas, and spatial variation in predation (in part in collaboration with middle school teachers and students in “The Moonsnail Project”).
I enjoy collaborating on my research with students at all levels, from undergraduate to PhD. Recent and current work in collaboration with graduate students includes: the ecological response of the Iceland molluscan fauna to the invasion of predatory gastropods during the Pliocene; taphonomy of an ophiuroid mass mortality bed in Mexico; geographic variation in drilling predation in Brazil and Argentina; relationship between drilling predation and prey diversity in the US Coastal Plain and Europe; conditions under which cannibalism by drilling predators occurs; metabolism and extinction susceptibility in Plio-Pleistocene bivalves. In addition, I serve as principal investigator on the National Science Foundation project: “Research Experiences for Undergraduates in Biodiversity Conservation” (Greg Dietl, Paleontological Research Institution, co-PI). This program has brought nine students per year from across the US to UNCWduring summers 2008-1010 to collect and compare data from the fossil and archeological record and modern marine communities to understand anthropogenic ecosystem change.
GLY 337 Invertebrate Paleontology
GLY 535 Stratigraphic Paleontology
College of Wooster
B.A., Geology 1975
Harvard University A.M., Geology 1977
Harvard University Ph.D., Geology 1979
Phi Beta Kappa, June 1974
Sigma Xi, June 1975
Outstanding Faculty Member of the School of Engineering, University of Mississippi, 1989-1990
University of North Dakota Sigma Xi Faculty Award for Outstanding Scientific Research, 1995
Elected Fellow of the Geological Society of America, 1995
President, Paleontological Society, 2000-2002
Association for Women Geoscientists Outstanding Educator Award, 2003
Elected Fellow of the American Association for the Advancement of Science, 2004
President, Board of Trustees, Paleontological Research Institution, 2004-2006
University of North Carolina Wilmington Award for Faculty Scholarship, 2005
Elected Centennial Fellow of the Paleontological Society, 2006
Kelley, P.H., J. Bryan, and T.A. Hansen (eds). 1999. The Evolution-Creationism Controversy II:Perspectives on Science, Religion, and Geological Education. The Paleontological Society Papers, vol. 5 (series editor, W. Manger).
Kowalewski, M., and P.H. Kelley (eds). 2002.The Fossil Record of Predation.The Paleontological Society Papers, vol. 8, 398 p.
Kelley, P.H., M. Kowalewski, and T.A. Hansen (eds). 2003.Predator-Prey Interactions in the Fossil Record.Kluwer Academic/Plenum Press, 464 p.Kelley, P.H., and R.K. Bambach (eds). 2008.From Evolution to Geobiology: Paleontology at the Start of a New Century.Paleontological Society Papers, 374 p.
Kelley, P.H. 2000. Studying evolution and keeping the faith. Geotimes 45(12):22-23,41.
Kelley, P.H., and T.A. Hansen. 2001. The role of ecological interactions in the evolution of naticid gastropods and their molluscan prey. Pp. 149-170. In W. Allmon and D. Bottjer, Evolutionary Paleoecology. Columbia University Press, New York.
Kelley, P.H., and T.A. Hansen. 2001. The Mesozoic marine revolution, p. 94-97. In D.E.G. Briggs and P.R. Crowther, Palaeobiology II. Blackwell Science, Oxford.
Kelley, P.H., T.A. Hansen, S.E. Graham, and A.G. Huntoon. 2001. Temporal patterns in the efficiency of naticid gastropod predators during the Cretaceous and Cenozoic of the United States Coastal Plain. Palaeogeography, Palaeoclimatology, Palaeoecology 166(1/2):165-176.
Kelley, P.H. and T.A. Hansen. 2001. Extinction. New Book of Knowledge, 5:425-426. (also published online at http://go.grolier.com:80/)
Dietl, G.P. and P.H. Kelley. 2001. Mid-Paleozoic latitudinal predation gradient: Distribution of brachiopod ornamentation reflects shifting Carboniferous climate. Geology 29:111–114.
Dietl, G.P., P.H. Kelley, R. Barrick and W. Showers. 2002. Escalation and extinction selectivity: morphology versus isotopic reconstruction of bivalve metabolism. Evolution 56(2):284-291.
Hansen, T.A., and P.H. Kelley. 2002. Moonsnail project: roles of technology and research in learning science. pp. 57-64, In Chambers, J.A. (ed.) Selected Papers from the Thirteenth International Conference on College Teaching and Learning. The Center for the Advancement of Teaching and Learning.
Dietl, G. P. and P.H. Kelley. 2002. The fossil record of predator-prey arms races: coevolution and escalation hypotheses, p. 353-374. In M. Kowalewski and P.H. Kelley (eds). The Fossil Record of Predation. The Paleontological Society Papers, vol. 8.
Kelley, P.H., and T.A. Hansen. 2003. The fossil record of drilling predation on bivalves and gastropods, p. 113-139. In P.H. Kelley, M. Kowalewski, and T.A. Hansen (eds), Predator-Prey Interactions in the Fossil Record. Kluwer Academic/Plenum Press.
Hansen, T.A., P.H. Kelley, and J.C. Hall. 2003. Moonsnail Project: a scientific collaboration with middle school teachers and students. Journal of Geoscience Education 50(1):35-38.
Hansen, T.A., P.H. Kelley, and D.M. Haasl. 2004. Paleoecological patterns in molluscan extinctions and recoveries: Comparison of the Cretaceous-Tertiary and Eocene-Oligocene extinctions in North America. Palaeogeography, Palaeoclimatology, Palaeoecology 214(3):233-242.
Reinhold, M.E., and P.H. Kelley. 2005. The influence of anti-predatory morphology on survivorship of the Owl Creek Formation molluscan fauna through the end-Cretaceous extinction. Palaeogeography, Palaeoclimatology, Palaeoecology 217:143-153.
Kelley, P.H., and T.A. Hansen. 2006. Comparisons of class- and lower taxon-level patterns in naticid gastropod predation, Cretaceous to Pleistocene of the U.S. Coastal Plain. Palaeogeography, Palaeoclimatology, Palaeoecology 236(3/4):302-320.
Dietl, G.P., and P.H. Kelley. 2006. Can naticid gastropod predators be discriminated by the holes they drill? Ichnos 13:1-6.
Kelley, P.H., and T.A. Hansen. 2007. A case for cannibalism: Confamilial and conspecific predation by naticid gastropods, Cretaceous through Pleistocene of the United States Coastal Plain, p. 151-170. In Elewa, A.M.T. (ed.), Predation in Organisms: A Distinct Phenomenon. Springer Verlag.
Kelley, P.H., and T.A. Hansen. 2007. Latitudinal patterns in naticid gastropod predation along the east coast of the United States: a modern baseline for interpreting temporal patterns in the fossil record, p. 284-299. In: Bromley, R.G., Buatois, L.A., Mángano, M.G., Genise, J.F., and Melchor, R.N. (eds.), Sediment-Organism Interactions: A multifaceted Ichnology. SEPM Special Publications, v. 88.
Kelley, P.H. 2008. Role of bioerosion in taphonomy: effect of predatory drillholes on preservation of mollusc shells, p. 451-470. In Tapanila, L., and M. Wisshak (eds), Current Developments in Bioerosion. Springer.
Kelley, P.H., 2008. The View from the Top: A Century of PS Presidents’ Perspectives on the Paleontological Society and Paleontology, p. 1-15. In Kelley, P.H., and R.K. Bambach (eds), From Evolution to Geobiology: Paleontology at the Start of a New Century. Paleontological Society Papers.
Bruce, K., J. Horan, P.H. Kelley, and M. Galizio. 2009. Teaching Evolution in the Galapagos. Journal of Effective Teaching 9(2):13-28.
Kelley, P.H. 2009. Stephen Jay Gould’s Winnowing Fork: Science, Religion, and Creationism, p. 171-188. In Allmon, W.D., P.H. Kelley, and R.D. Ross (eds), Stephen Jay Gould: Reflections on His view of Life. Oxford University Press.
Kelley, P.H. 2009. Teaching Evolution During the Week and Bible Study on Sunday: Perspectives on Science, Religion, and Intelligent Design, P. 163-179. In Schneiderman, J.S., and W.D. Allmon (eds), For the Rock Record: Geologists On Intelligent Design Creationism. University of California Press.
Kelley, P.H. 2009. A College Honors Seminar on Evolution and Intelligent Design: Successes and Challenges. Journal of Effective Teaching 9(2):29-37.
Dietl, G., S. Durham, and P. Kelley. 2010. Shell repair as a reliable indicator of bivalve predation by shell-wedging gastropods in the fossil record. Palaeogeography, Palaeoclimatology, Palaeoecology 296:174-184.