Biology & Marine Biology

Faculty & Staff

Darin S. Penneys, Assistant Professor

PenneysPh.D., Botany, University of Florida, Gainesville, FL, 2007
M.S., Botany, University of Florida, Gainesville, FL, 2001
B.S., Biology, Southern Connecticut State University, New Haven, CT, 1992
Dobo Hall 224 | (910) 962-3046 | 601 South College Road, Wilmington, NC 28403-5915 |

Research Interests: My primary interests are in the systematics, natural history, and evolution of angiosperms. Current projects include phylogenetic studies of Melastomataceae, using a global, multi-locus, molecular phylogeny that will serve as a foundation for a revised familial classification and biogeographical analyses. The Melastomataceae are one of the most diverse families of angiosperms and as such serve as excellent models for small- and large-scale investigations. My research combines fieldwork, herbarium, and molecular methods for studying speciation and adaptive radiations, plant-animal interactions pertaining to pollinator shifts and reward systems, ant- and mite-plant mutualisms, morphological character evolution, conservation, and phylogeography.


33. Penneys, D.S., C. Ulloa Ulloa, D.A. Neill, and D. Fernández. 2015. A new species of Chalybea (Blakeeae, Melastomataceae) from the Ecuador-Peru border. Phytokeys. Submitted.

32. Michelangeli, F.A., A.N. Nicolas, M. Reginato, G. Ocampo, R. Goldenberg, F. Almeda, W.S. Judd, E.R. Becquer-Granados, J.D. Skean, Jr., R. Kriebel, K. Sosa, M.K. Caddah, G. Ionta, D.S. Penneys, M. Alvear, J.M. Burke, and L. Majure. Historical Biogeography of the Neotropical tribe Miconieae (Melastomataceae) reveals a pattern of progressive colonization from Amazonia to other Neotropical forests. Proceedings of the National Academy of Sciences. Submitted.

31. Almeda, F., M. Alvear, D.S. Penneys, and F.A. Michelangeli. Melastomataceae. In: R. Bernal (ed.). Catalogue of the plants of Colombia. In press.

30. Morales-Puentes, M.E., D.S. Penneys, W.S. Judd, and F. González. Revisión taxonómica del clado Chalybea/Huilaea (Melastomataceae: Blakeeae). Revista de la Academia Colombiana de Ciencias Exactas, Fisicas y Naturales. In press.

29. Dellinger, A.S., D.S. Penneys, Y.M. Staedler, L. Fragner, W. Weckwerth, and J. Schoenenberger. 2014. A specialized bird pollination system with a bellows mechanism for pollen transfer and staminal food body rewards. Current Biology 24: 1615-1619. [doi: 10.1016/j.cub.2014.05.056]

28. Almeda, F., and D.S. Penneys. 2014. New and reconsidered species of Tropical American Melastomataceae. Brittonia 66: 160-169. [doi:10.1007/s12228-013-9320-2]

27. Penneys, D.S. 2013. Preliminary phylogeny of the Astronieae (Melastomataceae) based on nuclear and plastid DNA sequence data, with comments on the Philippine endemic genus, Astrocalyx. Philippine Journal of Science 142 (Special Issue): 159-168.

26. Penneys, D.S., and W.S. Judd. 2013. Combined molecular and morphological phylogenetic analyses of the Blakeeae (Melastomataceae). International Journal of Plant Sciences 174: 802-817. [doi:10.1086/670011]

25. Penneys, D.S., and W.S. Judd. 2013. New combinations and a revised circumscription for the Blakeeae (Melastomataceae). PhytoKeys 20: 17-32. [doi:10.3897/phytokeys.20.4344]

24. Michelangeli, F.A., P.J.F. Guimaraes, D.S. Penneys, F. Almeda, and R. Kriebel. 2013. Phylogenetic relationships and Distribution of New World Melastomeae (Melastomataceae). Botanical Journal of the Linnean Society 171: 38-60. [doi:10.1111/j.1095-8339.2012.01295.x]

23. Renner, S.S., D. Triebel, F. Almeda, R. Goldenberg, H. Mendoza-Cifuentes, F.A. Michelangeli, D.S. Penneys, R.D. Stone, and C. Ulloa. 2012. MELnet: Melastomataceae.Net (version 1.0, May 2011). In: F. Bisby, Y. Roskov, A. Culham, T. Orrell, D. Nicolson, L. Paglinawan, N. Bailly, W. Appeltans, P. Kirk, T. Bourgoin, G. Baillargeon, D. Ouvrard (eds). Species 2000 & ITIS Catalogue of Life, 30th April 2012. Digital resource at Species 2000: Reading, UK.

22. Penneys, D.S., and E. Cotton. 2011. Melastomataceae. Pp. 393-423. In: R. Valencia, N. Pitman, S. León-Yánez, and P.M. Jørgensen (eds). Libro rojo de las plantas endémicas del Ecuador, II Edition. Publicaciones del Herbario QCA, Pontifícia Universidad Católica del Ecuador, Quito.

21. Judd, W.S., G.M. Ionta, J.D. Skean, Jr., K. Campbell, and D.S. Penneys. 2011. Noteworthy collections of Heterotis rotundifolia in Jamaica and Dominica. Castanea 76: 311-312.

20. Wanntorp, L., C. Puglisi, D.S. Penneys, and L. Ronse De Craene. 2011. Multiplications of floral organs in flowers - a case study in Conostegia (Melastomataceae, Myrtales). Pp. 218-235. In: L. Wanntorp and L. Ronse De Craene (eds). Flowers on the Tree of Life. Systematics Association Special Volume, Cambridge University Press, Cambridge, England.

19. Penneys, D.S., and W.S. Judd. 2011. Phylogenetics and morphology in the Blakeeae (Melastomataceae). International Journal of Plant Sciences 172: 78-106. [doi:10.1086/657284]

18. Penneys, D.S., W.S. Judd, F.A. Michelangeli, and F. Almeda. 2010. Henrietteeae (Melastomataceae): A new Neotropical berry-fruited tribe. Systematic Botany 35: 783-800. [doi:10.1600/036364410X539862]

17. Morales-Puentes, M.E., and D.S. Penneys. 2010. New species of Chalybea Naud. and Huilaea Wurdack. Brittonia 62: 26-34. [doi:10.1007/s12228-009-9092-x]

16. Penneys, D.S., and L. Jost. 2009. Blakea attenboroughii (Melastomataceae: Blakeeae): A new species from Ecuador. Proceedings of the California Academy of Sciences 60: 69-72.

15. Goldenberg, R., D.S. Penneys, F. Almeda, W.S. Judd, and F.A. Michelangeli. 2008. Phylogeny of Miconia (Melastomataceae): Patterns of stamen diversification in a megadiverse Neotropical genus. International Journal of Plant Science 169: 963-979. [doi:10.1086/589697]

14. Judd, W.S., J.D. Skean, Jr., D.S. Penneys, and F.A. Michelangeli. 2008. A new species of Henriettea (Melastomataceae) from the Sierra de Baoruco, the Domincan Republic. Brittonia 60: 217-227. [doi:10.1007/s12228-008-9021-4]

13. Varassin, I. G., D.S. Penneys, and F.A. Michelangeli. 2008. Comparative anatomy and morphology of nectar-producing Melastomataceae. Annals of Botany 102: 899-909. [doi:10.1093/aob/mcn180]

12. Bécquer-G., E. R., K.M. Neubig, W.S. Judd, F.A. Michelangeli, J.R. Abbott, and D.S. Penneys. 2008. Preliminary molecular phylogenetic studies in Pachyanthus (Miconieae, Melastomataceae). Botanical Review 74: 37-52. [doi:10.1007/s12229-008-9010-z]

11. Michelangeli F.A., W.S. Judd, D.S. Penneys, J.D. Skean, Jr., E.R. Bécquer-G., R. Goldenberg, and C.V. Martin. 2008. Multiple events of dispersal and radiation of the tribe Miconieae (Melastomataceae) in the Caribbean. Botanical Review 74: 53-77. [doi:10.1007/s12229-008-9004-x]

10. Almeda, F., P.E. Berry, A. Freire-Fierro, A. Gröger, B.K. Holst, N.G. Luckana, F.A. Michelangeli, T. Morley, D.S. Penneys, S.S. Renner, O.R. Robinson, J.J. Wurdack, and K. Yatskievych. 2007. Melastomataceae. Pp. 397-417. In: V. Funk, T. Hollowell, P. Berry, C. Kelloff, and S.N. Alexander (eds.). Checklist of the plants of the Guiana Shield (Venezuela: Amazonas, Bolivar, Delta Amacuro; Guyana, Surinam, French Guiana). Contributions from the United States National Herbarium, Washington D.C.

9. Blanco, M.A., W.M. Whitten, D.S. Penneys, N.H. Williams, K.M. Neubig, and L. Endara. 2006. A simple and safe method for rapid drying of plant specimens using forced-air space heaters. Selbyana 27: 83-87.

8. Penneys, D.S., and W.S. Judd. 2005. A cladistic analysis and systematic revision of Charianthus (Miconieae: Melastomataceae) using morphological and molecular characters. Systematic Botany 30: 559-584. [doi:10.1600/0363644054782125]

7. Judd, W.S., and D.S. Penneys. 2004. Taxonomic studies in the Miconieae (Melastomataceae). VIII. A revision of the species of the Miconia desportesii complex on Hispaniola. Rhodora 106: 124-147.

6. Michelangeli, F.A., D.S. Penneys, J. Giza, D.E. Soltis, M.H. Hils, and J.D. Skean, Jr. 2004. A preliminary phylogeny of the tribe Miconieae (Melastomataceae) based on nrITS sequence data and its implications on inflorescence position. Taxon 53: 279-290.

5. Penneys, D.S., and W.S. Judd. 2004. Two new species of Charianthus (Melastomataceae: Miconieae) from the Lesser Antilles. Brittonia 56: 19-27. [doi:10.1663/0007-196X(2004)056[0151:TNSOCM]2.0.CO;2]

4. Judd, W.S., D.S. Penneys, and J.D. Skean, Jr. 2004. Rediscovery of Ossaea alloeotricha, an endemic of the high-elevation Massif de la Hotte, Haiti, and its transfer to Miconia (Melastomataceae: Miconieae). Brittonia 56: 56-62. [doi:10.1663/0007-196X(2004)056[0159:ROOAAE]2.0.CO;2]

3. Penneys, D.S., and W.S. Judd. 2003. The resurrection and lectotypification of Tetrazygia fadyenii (Melastomataceae: Miconieae): A hummingbird-pollinated treelet endemic to Jamaica. Sida 20: 877-884.

2. Penneys, D.S. 2001 - onward. Melastomataceae of the World [online]. Available at:

1. Penneys, D.S. 1996. A guide to the common flowering plant families of Monteverde, Costa Rica. Published by the author. 65 pp.